Smith-Paredes, D., Núñez-León, D., Soto-Acuña, S., O’Connor, J., Botelho, J. F., & Vargas, A. O. (2018). Dinosaur ossification centres in embryonic birds uncover developmental evolution of the skull. Nature Ecology & Evolution. Dec;2(12):1966-1973. doi: 10.1038/s41559-018-0713-1. Epub 2018 Nov 19.

Vargas AO, Ruiz-Flores M, Soto-Acuña S Haidr N, Acosta-Hospitaleche C, Ossa-Fuentes L and Muñoz-Walther V. 2017. The origin and evolutionary consequences of skeletal traits shaped by embryonic muscular activity, from basal theropods to modern birds. Integrative and Comparative Biology. DOI: 10.1093/icb/icx074 (Aceptado, en prensa)

Chimento NR, Agnolin FL, Novas FE, Ezcurra MD, Salgado L,  Isasi MP, Suárez M, De La Cruz R, Rubilar-Rogers D, Vargas AO. 2017.  Forelimb posture in Chilesaurus diegosuarezi (Dinosauria, Theropoda) and its behavioral and phylogenetic implications. Ameghiniana doi:10.5710/AMGH.11.06.2017.3088

Vargas AO, Krabichler Q and Guerrero-Bosagna C. 2016. An epigenetic perspective on the midwife toad experiments of Paul Kammerer (1880-1926). J. Exp. Zool. (Mol. Dev. Evol.) 00B:114 PDF

Botelho JF, Smith-Paredes D, Soto-Acuña S, Núñez-León D, Palma V, Vargas AO. 2016. Greater growth of proximal metatarsals in bird embryos and the evolution of hallux position in the grasping foot. J. Exp. Zool. (Mol. Dev. Evol.) 00B:113PDF

Botelho, J. F., Smith-Paredes, D., Soto-Acuña, S., O’Connor, J., Palma, V. and Vargas, A. O. (2016), Molecular development of fibular reduction in birds and its evolution from dinosaurs. Evolution.

Ossa-Fuentes  L, Mpodozis J, Vargas AO.  2015. Bird embryos uncover homology and evolution of the dinosaur ankle. Nature Communications 6, 8902.

Soto-Acuña S, Rubilar-Rogers D, Otero R, Vargas AO. Arcosaurios no avianos de Chile. En: Vertebrados Fósiles de Chile. Publicación Ocasional del Museo Nacional de Historia Natural, Chile, 63, Editores: David Rubilar-Rogers, Rodrigo A. Otero, Alexander Vargas M., Michel Sallaberry, pp.209-263

Botelho JF,  Smith-Paredes D y Vargas AO. 2015.  Altriciality and the Evolution of Toe Orientation in Birds. Evolutionary Biology. DOI 10.1007/s11692-015-9334-7. Published online 29 June 2015. PDF

Salgado L, Novas F,  Suárez M, de la Cruz R, Isasi M, Rubilar-Rogers D  y  Vargas AO. 2015. Upper jurassic sauropods in the Chilean patagonia. Ameghiniana DOI:10.5710/AMGH.07.05.2015.2883

Botelho, JF; Smith-Paredes, D; Soto-Acuña, S; Mpodozis, J; Palma, V; Vargas, AO. (2015) Skeletal plasticity in response to embryonic muscular activity underlies the development and evolution of the perching digit of birds. Scientific Reports Online 14/05/15.

Novas, Fernando; Salgado, Leonardo; Suárez, Manuel; Agnolín, Federico; Ezcurra, Martín; Chimento, Nicolás; de la Cruz, Rita; Isasi, Marcelo; Vargas, Alexander O. y Rubilar-Rogers, David. (2015) An enigmatic plant-eating theropod from the Late Jurassic period of Chile. Nature. (Publicado online el 27 de abril de 2015)

Vargas AO (2015) Comentario del libro “Flying Dinosaurs: How Fearsome Reptiles Became Birds” de Pickerel, J. Journal of Field Ornithology.

Vargas AO (2015) Comentario del libro “Avian Ancestors: A Review of the Phylogenetic Relationships of the Theropods Unenlagiidae, Microraptoria, Anchiornis and Scansoriopterygidae” de Agnolin F y Novas F. Hornero 30(1): 33-34.

Vargas A.O., Rubilar-Rogers D. y Soto-Acuña, S. (2015) La Evolución de los Ancestros de las Aves. In: Aves Rapaces de Chile, 2ª edición. Muñoz A., J. Rau, and J. Yañez, eds. Editorial CEA (en prensa)

Botelho JF, Ossa-Fuentes L, Soto-Acuña S, Smith-Paredes D, Nuñez-León D, et al. (2014) New Developmental Evidence Clarifies the Evolution of Wrist Bones in the Dinosaur–Bird Transition. PLoS Biol 12(9): e1001957. doi:10.1371/journal.pbio.1001957 Link: Synopsis:

Botelho J, Smith-Paredes D, Soto-Acuña S, Nuñez-Leon D,Vargas AO. 2014. The Developmental Origin of Zygodactyl Feet and its Inferred Loss in the Evolution of Passeriformes. Proceedings of the Royal Society B (Accepted May 30th). PDF

Otero, R.A., Soto-Acuña, S.,Vargas, A.O., Rubilar-Rogers, D. A new postcranial skeleton of an elasmosaurid plesiosaur from the Upper Cretaceous of central Chile and reassessment of the historic species Cimoliasaurus andium Deecke.  Cretaceous Research.

Otero RA, Soto-Acuña S, Gutstein C., Vargas AO, Rubilar-Rogers D., Yury-Yáñez R., Bastías J., Ramírez C. 2014. New chondrichthyans from the Late Cretaceous (Campanian–Maastrichtian) of Marambio and James Ross islands, Antarctica. Journal of Paleontology 88: 411-420

Salinas-Saavedra MGonzalez-Cabrera COssa-Fuentes L Botelho JFRuiz-Flores M y  Vargas AO. 2014. New developmental evidence supports a homeotic frameshift of digit identity in the evolution of the bird wing. Frontiers in Zoology 11:33. PDF

Otero RA, Soto-Acuña S, Vargas AO, Rubilar-Rogers D., Yury-Yáñez R., Gutstein C. 2013. Additions to the diversity of Elasmosaurid plesiosaurs from the upper cretaceous of Antartica. Gondwana Research. Available online 12 August 2013,

Vargas A.O. 2012.  El destino de las ideas sobre evolución humana de Max Westenhofer (1871-1957), profesor de la Universidad de Chile. Libro  “Darwin y la Evolución: Avances en la universidad de Chile” . Editorial Universitaria.

Otero R.A., D. Rubilar-Rogers, R. Yury-Yáñez R, A.O. Vargas,  F.A. Mourgues y C.S. Gutstein. 2012. A new species of chimaeriform (Chondrichthyes; Holocephali) from the uppermost Cretaceous of the López de Bertodano Formation, Seymour Island, Antarctica. Antarctic Science Firstview 1-8. PDF

Rubilar-Rogers D., R.A. Otero, R.Y. Yáñez, A.O. Vargas,  y C.S. Gutstein. 2012. An overview of the dinosaur fossil record from Chile. Journal of South American Earth Sciences 37:  242–255. PDF

Salinas-Saavedra M. y A.O. Vargas. 2011. Cortical Cytasters: A highly conserved developmental trait of Bilateria with similarities to Ctenophora.  EvoDevo 2: 23. PDF

 Kellner A.W.A, D. Rubilar-Rogers, A.O. Vargas y M. Suárez.  2011. A new titanosaur sauropod from the Atacama Desert, Chile. Anais da Academia Brasileira de Ciências 83(1): 211-219. PDF

Young RL, V Caputo, M Giovannotti, T Kohlsdorf, AO Vargas, GE May, y GP Wagner. 2009. Evolution of digit identity in the three-toed Italian skink Chalcides chalcides: a new case of Digit Identity Frame Shift. Evolution & Development 11 (6): 647-658 PDF

Vargas AO, GP Wagner y JA Gauthier. 2009. Limusaurus and bird digit identity. Available from Nature Precedings (archivo citable, no es “peer-reviewed”) <;  PDF . 

Vargas AO. 2009. Did Paul Kammerer discover epigenetic inheritance? A modern look at the controversial midwife toad experiments. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution. 312B: 667-678. PDF. Editorial por G. Wagner PDF. Comentario en Science por E. Pennissi PDF

Vargas AO, Fallon JF, Rubilar-Rogers D, Mpodozis J. 2009.  The evolution and development of the perching digit of birds. I Simposio-Paleontología en Chile, 2-3 Oct. 2008. Libro de Actas, pp. 174-176. PDF

Gutstein C.S., Cozzuol M.A., Vargas A.O., Suárez M.E., Schultz, C.L. and Rubilar-Rogers, D.  2009. Patterns of skull variation of Brachydelphis (Cetacea, Odontoceti) from the neogene of the southeastern pacific. Journal of Mammalogy 90 (2):504-519. PDF

Vargas A.O. and G.P. Wagner. 2009 Frame-shifts of digit identity in bird evolution and Cyclopamine-treated wings. Evolution & Development 11(2): 163-189 PDF

Vargas A.O., T. Kohlsdorf, J.F. Fallon, J. Vandenbrooks, and G.P. Wagner. 2008. The evolution of HoxD-11 expression in the bird wing: insights from Alligator mississippiensis. PLoS ONE  3(10): e3325. doi:10.1371/journal.pone.0003325 PDF

Wagner G.P. and A.O. Vargas. 2008 On the nature of thumbs. Genome Biology 9(3):213 PDF

Vargas A.O. 2005 Beyond Selection. Revista Chilena de Historia Natural, 2005. 78 (4): 739-752 PDF

Vargas A.O. and J.F. Fallon. 2005 The digits of the wing of birds are 1, 2 and 3. A review. Journal of Experimental Zoology part B: Molecular and Developmental Evolution 304 (3):206-219 PDF

Vargas A.O. and F. Aboitiz.  2005 How ancient is the adult swimming capacity in the lineage leading to euchordates?. Evolution & Development.  7:171-174 PDF

Vargas A.O. and J.F. Fallon. 2005 Birds have dinosaur wings: The molecular evidence. Journal of Experimental Zoology part B: Molecular and Developmental Evolution 304 (1): 86-90 PDF. Comentario en Science por E. Pennissi PDF

Cecchi, C., A. Vargas, C. Villagra, and J. Mpodozis. 2004  Answering Cuvier: notes on the systemic/historic nature of living beings. Cybernetics and Human Knowing 11: p. 11 – 19 PDF

Vargas A., and D. Rubilar-Rogers. 2004 La Evolución de los Ancestros de las Aves. In: Aves Rapaces de Chile. Muñoz A., J. Rau, and J. Yañez, eds. Editorial CEA pp. 107-119 PDF

Chiappe, L. M., and A. Vargas. 2003 Emplumando dinosaurios: La transición evolutiva de terópodos a aves. Hornero 18: 1-11. PDF

Vargas A. 1999 The evolution of Arm Size in Theropod Dinosaurs: A Developmental Hypothesis. Noticiario Mensual MNHN Chile. 338:16-19 PDF

Mayor, R., R. Young, and A. Vargas.  1999 Development of Neural Crest in Xenopus. Current Topics in Developmental Biology 41: 85-114 PDF